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Study of biology at TH Darmstadt; doctoral thesis on “Queen Polymorphism in the North American ant Leptothorax sp. A”, supervised by Prof. A. Buschinger and supported by German Academic Scholarship Foundation; ?doctoral examination in May 1988.

June 1988 to June 1989 post-doc in the lab of Prof. E.O. Wilson, Harvard University, Cambridge, Mass. Studies on the systematics, population biology, and ecology of North American ants of the genus Leptothorax; field work in Quebec, New England, Alberta and British Columbia.

September 1989 – Februar 1995 scientific assistant at the chair of Prof. Dr. B. H?lldobler, University of Würzburg; establishment of the research areas population biology and sociogenetics; studies on reproductive conflict and conflict resolution in ant societies. Field studies in North America, India, Brazil, Turkey.?

July 1994 Habilitation in zoology with a thesis on "Cooperation and Conflict in Ant Societies",?

March 1995 – September 1996 Heisenberg Fellowship

October 1996 – March 2000 Professor of Zoology at University of Erlangen-Nürnberg?

Since April 2000 Professor of Zoology at University of Regensburg?
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For many years member of senate and joint committee of German Research Foundation, senate committee on Research Training Groups, and the Excellence Initiative Grants Committee.

Research topics

Conflict and conflict resolution in insect societies

Evolution of alternative reproductive tactics and life histories

Reproduction, mating, and aging in social insects